The characteristic of the lion as the "king of the jungle" goes back to the influence of The Physiologus, an early Christian book about animal symbolism which spread into many cultures and generally had great influence in Western culture. First written in Greek in the second century AD, the book was translated into Latin in about 400 AD, next into Ethiopic and Syriac, then into many European and Middle-Eastern languages. Many illuminated manuscript copies such as the Bern Physiologus survive. It retained its influence over ideas of the "meaning" of animals in Europe for over a thousand years. It was a predecessor of bestiaries (books of beasts). Medieval poetical literature is full of allusions that can be traced to the Physiologus tradition; the text also exerted great influence on the symbolism of medieval ecclesiastical art.